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Investigations on the prior arts on two-tier femtocell networks indicate that the analysis fundamental capacity limit with optimal power and sub-channel adaptation is not well considered. Based on the motivation, mapping the conventional Interference Temperature Model (ITM) to the two-tier femtocell network, the problem model is formulated. Then, we derive a fundamental relation for optimal sub-channel and power adaptation and the closed-form expression for the achievable capacity. Numerical results confirm our mathematical derivation, which provide theoretical guidance for resource allocation of two-tier femtocell networks and other wireless hierarchical networks with frequency reuse.