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The significant advances of hardware manufacturing technology and the development of efficient software algorithms make technically and economically feasible a network composed of numerous, small, low-cost sensors using wireless communications, that is, a wireless sensor network. WSNs have attracted intensive interest from both academia and industry due to their wide application in civil and military scenarios. In hostile scenarios, it is very important to protect WSNs from malicious attacks. Due to various resource limitations and the salient features of a wireless sensor network, the security design for such networks is significantly challenging. In this article, we present a comprehensive survey of WSN security issues that were investigated by researchers in recent years and that shed light on future directions for WSN security.