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In randomly-deployed wireless mission critical networks, the crucial steps of ad hoc route setup and node localization are vulnerable to various security breaches and attacks. In this paper, we introduce SIRLoS, a lightweight secure integrated routing and localization scheme which addresses this problem by exploiting the security benefits of link directionality in directed networks that have found popularity for multimedia networking. SIRLoS is a circuit-based algorithm that leverages the resources of the base station and the hierarchical structure of the network to reconstruct the graph of the network, and detect any security violations in the neighborhood discovery and routing schemes. We demonstrate the performance of our algorithm, and provide security and attack analysis.